MSH6

Chr 2ADSomaticAR

mutS homolog 6

Also known as: GTBP, GTMBP, HNPCC5, HSAP, LYNCH5, MMRCS3, MSH-6, p160

The MSH6 protein heterodimerizes with MSH2 to form a mismatch recognition complex that binds to DNA mismatches and initiates post-replicative DNA mismatch repair. Mutations cause Lynch syndrome 5, familial endometrial cancer, and mismatch repair cancer syndrome 3 with autosomal dominant inheritance (though some cases show autosomal recessive inheritance). This gene is highly intolerant to loss-of-function variants (pLI near 1.0, LOEUF 0.498), indicating that complete loss of function is likely incompatible with normal development.

GeneReviewsOMIMResearchSummary from RefSeq, OMIM, UniProt
LOFmechanismAD/Somatic/ARLOEUF 0.503 OMIM phenotypes
Clinical SummaryMSH6
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Gene-Disease Validity (ClinGen)
mismatch repair cancer syndrome 1 · ARDefinitive

Definitive — sufficient evidence for diagnostic panels

3 total gene-disease associations curated

Population Constraint (gnomAD)
Constrained for loss-of-function variants (OE-LoF 0.34) despite low pLI — interpret in context.
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ClinVar Variants
248 unique Pathogenic / Likely Pathogenic· 409 VUS of 1200 total submissions
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Clinical Trials
12 active or recruiting trials — potential therapeutic options may be available
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GeneReview available — MSH6
Authoritative clinical overview · Recommended first read
Open GeneReview ↗

Population Genetics & Constraint

gnomAD v4 — loss-of-function & missense intolerance

Moderate LoF intolerance
LoF Constraint
0.50LOEUF
pLI 0.000
Z-score 4.51
OE 0.34 (0.230.50)
Moderately constrained

More LoF-intolerant than ~75% of genes

Missense Constraint
-2.78Z-score
OE missense 1.29 (1.231.37)
917 obs / 708.7 exp
Tolerant

Tolerant to missense variation

Observed / Expected Ratios
LoF OE0.34 (0.230.50)
00.351.4
Missense OE1.29 (1.231.37)
00.61.4
Synonymous OE1.50
01.21.6
LoF obs/exp: 18 / 53.6Missense obs/exp: 917 / 708.7Syn Z: -6.32
Curated Mechanism (G2P)Gene2Phenotype (DDG2P) ↗
definitiveMSH6-related Lynch syndromeLOFAD
definitiveMSH6-related constitutional mismatch repair deficiency syndromeLOFAR
DN
0.6260th %ile
GOF
0.4382th %ile
LOF
0.3452th %ile

This gene has evidence for multiple mechanisms of pathogenicity (loss-of-function and dominant-negative). The Badonyi & Marsh model scores dominant-negative highest among its predictions, but genomic evidence (constraint, ClinVar variant spectrum, and literature) most strongly supports loss-of-function (haploinsufficiency). Different variants in this gene may act through different mechanisms — interpret in context of the specific variant.

LOF1 literature citation · 90% of P/LP variants are LoF · LOEUF 0.50
DNprediction above median

Note: In-silico variant effect predictors (SIFT, PolyPhen, REVEL, CADD) may underestimate pathogenicity of missense variants in genes with GOF or DN mechanisms. Consider functional evidence and clinical context.

Literature Evidence

LOFGermline mutation of MSH6 as the cause of hereditary nonpolyposis colorectal cancerPMID:9354786

Predictions from Badonyi M, Marsh JA. PLoS ONE. 2024;19(8):e0307312. Mechanism ranking also informed by gnomAD constraint, ClinVar, and ClinGen data.

ClinVar Variant Classifications

1200 submitted variants in ClinVar

Classification Summary

Pathogenic163
Likely Pathogenic85
VUS409
Likely Benign292
Benign42
Conflicting196
163
Pathogenic
85
Likely Pathogenic
409
VUS
292
Likely Benign
42
Benign
196
Conflicting

Curated Variants Distribution

Classified variants from ClinVar · 5 ACMG categories

ClassificationLoFMissense + InframeNon-codingSynonymousTotal
Pathogenic
152
3
8
0
163
Likely Pathogenic
70
11
4
0
85
VUS
9
381
19
0
409
Likely Benign
1
11
79
201
292
Benign
1
5
6
30
42
Conflicting
196
Total2334111162311,187

LoF = frameshift, stop gained/lost, canonical splice · Counts from ClinVar esearch · Updated hourly

View in ClinVar →

Protein Context — Lollipop Plot

MSH6 · protein map & ClinVar variants

Showing all ClinVar variants across the protein. Search a specific variant to highlight its position.

3D Protein StructureAlphaFold

Clinical Trials

Active and recruiting trials from ClinicalTrials.gov

MMR MutationSmall Bowel AdenomaSmall-bowel Adenocarcinoma

Small Bowel Capsule Endoscopy in Lynch Syndrome

RECRUITING
NCT07472686Assistance Publique - Hôpitaux de ParisStarted 2025-06-16
Basal Cell Carcinoma of Skin, Site UnspecifiedEpidermoid CarcinomaLynch Syndrome

Determining the Prevalence of Muir-Torre Syndrome in Patients With Lynch Syndrome

NOT YET RECRUITING
NCT07201012Phase NACentre Hospitalier Universitaire de NīmesStarted 2025-12
Sampling of suspected skin lesions (in accordance with good care practices).Constitution of a biobank
Pancreatic CancerPancreatic Ductal AdenocarcinomaPDAC

A Prospective Registry for Patients at High-Risk for Pancreatic Cancer

RECRUITING
NCT06151223Mayo ClinicStarted 2021-07-13
Bio-specimen Collection: BloodBio-specimen Collection: Pancreatic JuiceMRI
Solid TumorAdvanced Solid TumorMetastatic Cancer

KPMNG Study of MOlecular Profiling Guided Therapy Based on Genomic Alterations in Advanced Solid Tumors II

RECRUITING
NCT05525858Seoul National University Bundang HospitalStarted 2022-09-28
AlectinibAtezolizumabErlotinib
Polycythemia VeraEssential ThrombocythaemiaMyelofibrosis

Prevalence Of Germline Gene Mutations In Patients With Myeloproliferative Neoplasms With Family History

NOT YET RECRUITING
NCT06923670Phase NAFondazione Policlinico Universitario Agostino Gemelli IRCCSStarted 2025-05-21
NGS testingNGS analysis for mutations in genes involved in familial predisposition to hematological malignancies
Uveal Melanoma

Identification of New Candidate Genes for Hereditary Predisposition to Uveal Melanoma

RECRUITING
NCT06550674Phase NACentre Jean PerrinStarted 2024-10-29
Constitutional exome analysis
Lynch SyndromeEndometrial Cancer

EC_ItaLynch: Mainstreaming the Diagnosis of Lynch Syndrome

NOT YET RECRUITING
NCT06501417Fondazione Policlinico Universitario Agostino Gemelli IRCCSStarted 2024-08-01
mainstreaming
BRCA1 MutationPOLD1 Gene MutationCDKN2A Mutation

An Intervention to Increase Genetic Testing in Families Who May Share a Gene Mutation Related to Cancer Risk and An Intervention to Help Patients and Their Primary Care Providers Stay Up-to-date About Uncertain Genetic Test Results

RECRUITING
NCT05420064Phase NAMemorial Sloan Kettering Cancer CenterStarted 2022-12-01
Intervention Arm At-risk Relative/ARR ContactsMyGene PortalStandard of Care
Lynch SyndromeLynch Syndrome ILynch Syndrome II

Videocapsule Endoscopy in Lynch Syndrome

RECRUITING
NCT05704010Phase NASan Raffaele UniversityStarted 2018-11-01
Video capsule endoscopy
Breast Cancer RiskOvarian Cancer RiskCancer Gene Mutation

Population Based Germline Testing for Early Detection and Prevention of Cancer

RECRUITING
NCT07498829Phase NAQueen Mary University of LondonStarted 2025-12-18
Genetic testing for Cancer Susceptibility Genes (CSGs) (BRCA1, BRCA2, RAD51C, RAD51D, BRIP1, PALB2, MLH1, MSH2, MSH6) and personalised breast and ovarian cancer risk
Deleterious BARD1 Gene MutationDeleterious BRCA1 Gene MutationDeleterious BRCA2 Gene Mutation

Surgery in Preventing Ovarian Cancer in Patients With Genetic Mutations

ACTIVE NOT RECRUITING
NCT02760849Phase NAM.D. Anderson Cancer CenterStarted 2016-05-02
Laboratory Biomarker AnalysisOophorectomyQuality-of-Life Assessment
Hereditary Breast/Ovarian Cancer (brca1, brca2)Lynch SyndromeGenetic Variation

Patient Centered Clinical Decision Support for Hereditary Cancer Syndromes

ENROLLING BY INVITATION
NCT06914726Phase NAHealthPartners InstituteStarted 2025-07-09
Patient Centered Clinical Decision Support (PC-CDS)
Clinical Literature
Open Research Assistant →
Key Publications
Landmark & review papers · by relevance
PubMed
Top 5 results · since 2015Search PubMed ↗