JAK2

Chr 9ADSomatic

Janus kinase 2

Also known as: JTK10

The JAK2 protein is a non-receptor tyrosine kinase that mediates cytokine and growth factor signaling by phosphorylating STAT transcription factors and plays a key role in hematopoiesis, immune responses, and chromatin regulation. Somatic mutations cause myeloproliferative disorders including polycythemia vera, myelofibrosis, acute myeloid leukemia, and thrombocythemia, while germline mutations cause autosomal dominant erythrocytosis. The gene is highly constrained against loss-of-function variation (LOEUF 0.341), reflecting its essential role in normal hematopoietic development.

GeneReviewsOMIMResearchSummary from RefSeq, OMIM, UniProt
MultiplemechanismAD/SomaticLOEUF 0.346 OMIM phenotypes
Clinical SummaryJAK2
Population Constraint (gnomAD)
Moderately constrained gene (pLI 0.65) — some intolerance to loss-of-function variants.
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Clinical Trials
12 active or recruiting trials — potential therapeutic options may be available
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GeneReview available — JAK2
Authoritative clinical overview · Recommended first read
Open GeneReview ↗

Population Genetics & Constraint

gnomAD v4 — loss-of-function & missense intolerance

LoF intolerant — likely haploinsufficient
LoF Constraint
0.34LOEUF
pLI 0.654
Z-score 5.66
OE 0.21 (0.140.34)
Highly constrained

Highly LoF-intolerant (top ~10% of genes)

Missense Constraint
0.90Z-score
OE missense 0.90 (0.830.96)
518 obs / 578.6 exp
Tolerant

Mild missense constraint

Observed / Expected Ratios
LoF OE0.21 (0.140.34)
00.351.4
Missense OE0.90 (0.830.96)
00.61.4
Synonymous OE1.11
01.21.6
LoF obs/exp: 13 / 60.6Missense obs/exp: 518 / 578.6Syn Z: -1.21
DN
0.5575th %ile
GOF
0.6345th %ile
LOF
0.4628th %ile

This gene has evidence for multiple mechanisms of pathogenicity (gain-of-function and loss-of-function). Both the Badonyi & Marsh prediction and the broader genomic evidence point to gain-of-function as the predominant mechanism. Different variants in this gene may act through different mechanisms — interpret in context of the specific variant.

GOFprediction above median · 1 literature citation
LOFLOEUF 0.34

Note: In-silico variant effect predictors (SIFT, PolyPhen, REVEL, CADD) may underestimate pathogenicity of missense variants in genes with GOF or DN mechanisms. Consider functional evidence and clinical context.

Literature Evidence

GOFThrombotic complications are a major cause of death in patients with Philadelphia chromosome-negative myeloproliferative neoplasms (MPNs), which are closely associated with the JAK2 V617F activating mutation.PMID:24858412

Predictions from Badonyi M, Marsh JA. PLoS ONE. 2024;19(8):e0307312. Mechanism ranking also informed by gnomAD constraint, ClinVar, and ClinGen data.

ClinVar Variant Classifications

0 submitted variants in ClinVar

Protein Context — Lollipop Plot

JAK2 · protein map & ClinVar variants

Showing all ClinVar variants across the protein. Search a specific variant to highlight its position.

3D Protein StructureAlphaFold

Clinical Trials

Active and recruiting trials from ClinicalTrials.gov

Aplastic AnemiaTPO Receptor Agonists

Standard Immunosuppressive Therapy Combined With Romiplostim N01 as First-line Treatment for Severe Aplastic Anemia

RECRUITING
NCT06613880Phase PHASE2Institute of Hematology & Blood Diseases Hospital, ChinaStarted 2024-10-24
standard IST combined with Romiplostim N01
Polycythemia VeraEssential ThrombocythaemiaMyelofibrosis

Prevalence Of Germline Gene Mutations In Patients With Myeloproliferative Neoplasms With Family History

NOT YET RECRUITING
NCT06923670Phase NAFondazione Policlinico Universitario Agostino Gemelli IRCCSStarted 2025-05-21
NGS testingNGS analysis for mutations in genes involved in familial predisposition to hematological malignancies
Giant Cell ArteritisTemporal ArteritisClonal Hematopoiesis of Indeterminate Potential

Clonal Hematopoiesis in Giant Cell Arteritis

NOT YET RECRUITING
NCT06244069ASST Fatebenefratelli SaccoStarted 2024-03
Temporal arterial biopsyWhole exome sequencingSingle cell transcriptomics
Myelofibrosis Transformation in Essential ThrombocythemiaPolycythemia Vera, Post-Polycythemic Myelofibrosis PhasePrimary Myelofibrosis

Elotuzumab for the Treatment of JAK2-Mutated Myelofibrosis

ACTIVE NOT RECRUITING
NCT04517851Phase PHASE2M.D. Anderson Cancer CenterStarted 2021-02-10
ElotuzumabQuestionnaire Administration
Insulin ResistanceObesity & OverweightEnergy Metabolism

Effects of Ketone Bodies on Insulin Sensitivity

RECRUITING
NCT07359625Phase NAUniversity of AarhusStarted 2025-12-02
Growth Hormone, HumanOral ketone supplement (KetoAid®, KE4)Saline infusion (placebo)
Myelofibrosis

Tasquinimod in Patients with Myelofibrosis Refractory to or Intolerant for JAK2 Inhibition

RECRUITING
NCT06605586Phase PHASE1, PHASE2Stichting Hemato-Oncologie voor Volwassenen NederlandStarted 2025-02-20
Tasquinimod
Myelofibrosis,MF

Pacritinib For Bone Marrow Fibrosis In Patients With Myelofibrosis Who Have Thrombocytopenia

RECRUITING
NCT07394153Phase PHASE2Grupo Español de Enfermedades Mieloproliferativas Crónicas PH NegativasStarted 2026-04-15
Pacritinib
Myeloproliferative DiseaseGermline Mutation

Unveiling the Germline Predisposition to Myeloproliferative Neoplasms

RECRUITING
NCT07204392Fondazione IRCCS Policlinico San Matteo di PaviaStarted 2022-06-27
T-cell LymphomaGraft Versus Host DiseaseLymphoma, T-Cell

Ruxolitinib Maintenance Post-Hematopoietic Stem Cell Transplant T-Cell Lymphoma

RECRUITING
NCT07356245Phase PHASE2Jonathan BrammerStarted 2026-02-12
RuxolitinibPositron emission tomography-computed tomographyBone Marrow Biopsy
Myelofibrosis

A Clinical Trial of TQ05105 Tablets Combined With TQB3617 Capsules in the Treatment of Myelofibrosis (MF)

RECRUITING
NCT06122831Phase PHASE1, PHASE2Chia Tai Tianqing Pharmaceutical Group Co., Ltd.Started 2023-12-12
TQ05105 TabletsTQB3617 Capsules
Myeloproliferative NeoplasmPolycythemia VeraEssential Thrombocythemia

Functional and Phenotypic Characterization of Monocytes in Myeloproliferative Syndromes

RECRUITING
NCT06361641Phase NAUniversity Hospital, AngersStarted 2024-05-29
Monocytes signatures in myeloproliferative neoplasms at diagnosis
E1 Ubiqutin-activating Enzyme, X-linked, Autoinflammatory, Somatic SyndromeVEXASVexas Syndrome

Pacritinib in Vacuoles, E1 Ubiqutin-activating Enzyme, X-linked, Autoinflammatory, Somatic (VEXAS) Syndrome

RECRUITING
NCT06538181Phase PHASE1Washington University School of MedicineStarted 2025-02-13
Pacritinib
Clinical Literature
Open Research Assistant →
Full-Text Mentions
NLP-detected gene mentions in article bodies · via PubTator3
PubTator3
Top 5 full-text resultsSearch PubTator3 ↗
Key Publications
Landmark & review papers · by relevance
PubMed
The MPL mutation.
Guglielmelli P et al.·Int Rev Cell Mol Biol
2021Review
Polycythaemia vera.
Harrison CN et al.·Nat Rev Dis Primers
2025Review
Plaque erosion risk and JAK2 V617F variant.
Wang S et al.·Eur Heart J
2025
Top 5 results · since 2015Search PubMed ↗