EIF2B5

Chr 3AR

eukaryotic translation initiation factor 2B subunit epsilon

Also known as: CACH, CLE, EIF-2B, EIF2Bepsilon, LVWM, VWM5

The protein is the epsilon subunit of eukaryotic translation initiation factor 2B (EIF2B), a GTP exchange factor that regulates protein synthesis by activating eukaryotic initiation factor 2. Mutations cause leukoencephalopathy with vanishing white matter, an autosomal recessive leukodystrophy that may be associated with ovarian failure in females. The pathogenic mechanism involves loss of function of the EIF2B complex, disrupting cellular protein synthesis regulation.

OMIMResearchSummary from RefSeq, OMIM, UniProt, Mechanism
LOFmechanismARLOEUF 0.441 OMIM phenotype
Clinical SummaryEIF2B5
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Gene-Disease Validity (ClinGen)
leukoencephalopathy with vanishing white matter 5 · ARDefinitive

Definitive — sufficient evidence for diagnostic panels

Population Constraint (gnomAD)
Constrained for loss-of-function variants (OE-LoF 0.26) despite low pLI — interpret in context.
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Clinical Trials
2 active or recruiting trials — potential therapeutic options may be available

Population Genetics & Constraint

gnomAD v4 — loss-of-function & missense intolerance

Moderate LoF intolerance
LoF Constraint
0.44LOEUF
pLI 0.064
Z-score 4.25
OE 0.26 (0.160.44)
Moderately constrained

More LoF-intolerant than ~75% of genes

Missense Constraint
0.78Z-score
OE missense 0.89 (0.810.97)
346 obs / 389.1 exp
Tolerant

Mild missense constraint

Observed / Expected Ratios
LoF OE0.26 (0.160.44)
00.351.4
Missense OE0.89 (0.810.97)
00.61.4
Synonymous OE0.95
01.21.6
LoF obs/exp: 10 / 38.5Missense obs/exp: 346 / 389.1Syn Z: 0.44
Curated Mechanism (G2P)Gene2Phenotype (DDG2P) ↗
definitiveEIF2B5-related leukoencephalopathy with vanishing white matterLOFAR

Predictions shown for reference only — model trained on dominant genes, not applicable to AR conditions.

DN
0.6356th %ile
GOF
0.5366th %ile
LOF
0.3357th %ile

The Badonyi & Marsh prediction model was trained exclusively on dominant disease genes. Predictions are not reliable for genes with autosomal recessive inheritance and are shown at reduced opacity for reference only.

Predictions from Badonyi M, Marsh JA. PLoS ONE. 2024;19(8):e0307312.

ClinVar Variant Classifications

0 submitted variants in ClinVar

Protein Context — Lollipop Plot

EIF2B5 · protein map & ClinVar variants

Showing all ClinVar variants across the protein. Search a specific variant to highlight its position.

3D Protein StructureAlphaFold
Clinical Literature
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